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In this case the only regularity to be noted is the rectangularity of
the completed surface. There are but few of the inner figures bordered by
four adjacent ones. The direct environment of the frog consists of two
figures; the guitar is hemmed in by three, the cock by five and the ostrich
(if that is what it really is) by six. The sum total can only be arrived by
careful counting.
It's of interest to us that only one figure is not a creature of some sort.
In our reading we haven't been able to find any stated meaning to the guitar,
but Escher did almost everything with a purpose.
Escher occasionally worked on irregular filling of plane surfaces such as
this print Mosaic II. He said they could never have been produced without
years of training in regular surface-filling and the only reason for their
existence is one's enjoyment of this difficult game.